A world classification of the Harmacloninae, a new subfamily of Tineidae (Lepidoptera: Tineoidea) / / Donald R. Davis.

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A world classification of the Harmacloninae, a new subfamily of Tineidae (Lepidoptera: Tineoidea) / / Donald R. Davis.

Davis, Donald R. (Donald Ray)

A world classification of the Harmacloninae, a new subfamily of Tineidae (Lepidoptera: Tineoidea) / / Donald R. Davis.

Washington, D.C. : Smithsonian Institution Press, 1998.

iii, 81 p. : ill., maps ; 28 cm.

Electronic Version:

Online Publication http://dx.doi.org/10.5479/si.00810282.597

Smithsonian contributions to zoology ; -- no. 597

The systematics, phylogeny, morphology, and distribution are summarized for the newly proposed subfamily Harmacloninae. Synapomorphies distinguishing this subfamily from the sister group, Myrmecozelinae, include (1) unique wing coupling system consisting of raised scales along the ventral hindmargin of the forewing interlocking into similar scales along the dorsal subcostal area of the hindwing; (2) pretarsus without arolium and pseudempodial seta; (3) the presence of paired abdominal tympanic organs; (4) enlargement of the sternal apophyses on the second abdominal sternum; and (5) aedoeagus with a basal, midventral keel. Cladistic analysis of the 22 species, using the genus Gerontha of the Myrmecozelinae as an outgroup, resulted in the recognition of two monophyletic genera: Micrerethista, with 10 of the 11 recognized species concentrated in the Oriental/Australian regions and a single species from equatorial Africa, and Harmaclona, consisting of 11 pantropical species. The following taxa are described as new: Micrerethista africana, M. bifida, M. denticulata, M. dissacca, M. fasciola, M. fusca, M. nigrapex, M. resima, Harmaclona afrotephrantha, H. hexacantha, H. robinsoni, H. tetracantha, and H. triacantha. Distribution maps and keys are provided for all species. Diagnostic characters of all taxa are fully illustrated by line drawings and photographs.The vicariant distributions of certain Harmacloninae suggest a minimum age of more than 90 m.y. for the subfamily. If tympanic organs in Harmacloninae evolved primarily as a defense against insectivorous, echolocating bats, as has been proposed for other Lepidoptera, then this would indicate that echolocating bats must have appeared earlier than the Harmacloninae—at least by the beginning of the late Cretaceous.

Also available electronically as a PDF file or files (requires Adobe Actobat Reader).

Bibliography Note:

Includes bibliographical references (p. 55-57).

Tineidae -- Classification.

Smithsonian contributions to zoology no. 597.

Catalog Source No.:

(OCoLC)ocm37546783

Horizon Information Portal 3.24_7488

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